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John’s Google Scholar page may be more up-to-date.


Mola, J. M., I. S. Pearse, M. L. Boone, E. Evans, M. J. Hepner, R. P. Jean, J. M. Kochanski, C. Nordmeyer, E. Runquist, T. A. Smith, J. P. Strange, J. Watson, and J. B. U. Koch. 2024. Range-wide genetic analysis of an endangered bumble bee (Bombus affinis, Hymenoptera: Apidae) reveals population structure, isolation by distance, and low colony abundance. Journal of Insect Science 24:19.


Janousek, W. M., M. R. Douglas, S. Cannings, M. A. Clément, C. M. Delphia, J. G. Everett, R. G. Hatfield, D. A. Keinath, J. B. U. Koch, L. M. McCabe, J. M. Mola, J. E. Ogilvie, I. Rangwala, L. L. Richardson, A. T. Rohde, J. P. Strange, L. M. Tronstad, and T. A. Graves. 2023. Recent and future declines of a historically widespread pollinator linked to climate, land cover, and pesticides. Proceedings of the National Academy of Sciences 120:e2211223120.


Otto, C. R. V., A. C. Schrage, L. L. Bailey, J. M. Mola, T. A. Smith, I. Pearse, S. Simanonok, and R. Grundel. 2022. Addressing Detection Uncertainty in Bombus affinis (Hymenoptera: Apidae) Surveys Can Improve Inferences Made From Monitoring. Environmental Entomology:nvac090.

Kendall, L.K., Mola, J.M., Portman, Z.M., Cariveau, D., Smith, H.G., Bartomeus, I. 2022. The potential and realized foraging movements of bees are differentially determined by body size and sociality. Ecology.

Montero-Castaño, A., J. B. U. Koch, T.-T. T. Lindsay, B. Love, J. M. Mola, K. Newman, and J. K. Sharkey. 2022. Pursuing best practices for minimizing wild bee captures to support biological research. Conservation Science and Practice. Early View Link

Rundlöf, M., C. Stuligross, A. Lindh, R. L. Malfi, K. Burns, J. M. Mola, S. Cibotti, and N. M. Williams. 2022. Flower plantings support wild bee reproduction and may also mitigate pesticide exposure effects. Journal of Applied Ecology. Early View Link


J.M. Mola, J. Hemberger, J. Kochanski, L. L. Richardson, and I. S. Pearse. 2021. Importance of forests in bumble bee biology and conservation. BioScience. Link

J.M. Mola, L. L. Richardson, G. Spyreas, D. N. Zaya, and I. S. Pearse. 2021. Long‐term surveys support declines in early season forest plants used by bumblebees. Journal of Applied Ecology:1365-2664.13886. Link

J.M. Mola, C. Stuligross, M.L. Page, D. Rutkowski, and N.M. Williams. 2021. Impact of “non-lethal” tarsal clipping on bumble bees (Bombus vosnesenskii) may depend on queen stage and worker size. Journal of Insect Conservation. 25:195–201. Open Access


Smith, T.A., J.P. Strange, E.C. Evans, B.M. Sadd, J.C. Steiner, J.M. Mola and K. Traylor-Holzer.(Eds.). 2020. Rusty Patched Bumble Bee, Bombus affinis, Ex Situ Assessment and Planning Workshop: Final Report. IUCN SSC Conservation Planning Specialist Group, Apple Valley, MN, USA

Mola J.M., M.R. Miller, S. O’Rourke and N.M. Williams. 2020. Wildfire reveals transient change to individual traits and population responses of a native bumble bee Bombus vosnesenskii. Journal of Animal Ecology, LINK

Mola J.M., M.R. Miller, S. O’Rourke & N.M. Williams. 2020. Forests do not limit bumble bee foraging movements in a montane meadow complex. Ecological Entomology, LINK

LoPresti, E.F., J. Goidell, J.M. Mola, M.L. Page, C.D. Specht, C. Stuligross, M.G. Weber, N.M. Williams, and R. Karban. 2020. A lever action hypothesis for pendulous hummingbird flowers: experimental evidence from a columbine, Annals of Botany 125(2): 59-65. DOI


Williams, N.M., J.M. Mola, Stuligross, C., Harrison, T., Page, M., Brennan, R., Rosenberger, N., & Rundlöf, M. 2019. Fantastic bees and where to find them: locating the cryptic overwintering queens of a western bumble bee. Ecosphere 9(1):e02056. Open Access

Mola, J. M. and N. M. Williams. 2019. A review of methods for the study of bumble bee movement. Apidologie 50: 497–514. DOI

Mola, J. M. 2019. Bumble bee movement ecology and response to wildfire with an emphasis on genetic mark-recapture techniques. Doctoral Dissertation. University of California Davis. LINK


LoPresti, E. F., J. I. V. Wyk, J. M. Mola, K. Toll, T. J. Miller, and N. M. Williams. 2018. Effects of wildfire on floral display size and pollinator community reduce outcrossing rate in a plant with a mixed mating system. American Journal of Botany 105:1154–1164.

Mola, J. M., and N. M. Williams. 2018. Fire-induced change in floral abundance, density, and phenology benefits bumble bee foragers. Ecosphere 9. Open Access


Mola, J. M., J. M. Varner, E. S. Jules, and T. Spector. 2014. Altered Community Flammability in Florida’s Apalachicola Ravines and Implications for the Persistence of the Endangered Conifer Torreya taxifolia. PLoS ONE 9:e103933. PDFKreye, J. K., J. M. Varner, J. K. Hiers, and J. Mola. 2013. Toward a mechanism for eastern North American forest mesophication: differential litter drying across 17 species. Ecological Applications 23:1976–1986.